The Rolls -Royce C Series engines were wet liner.
At the lower end, they were sealed with two O rings, often with Wellseal applied. Ar the top, the seal was metal to metal, again with Wellseal applied.
Cylinder liner protrusion was carefully controlled so that the liner provided a flame ring, to protect bthe head gasket.
The Bristol BVW engine sealed the liner at the lower end with two O rings. The block was drilled between the rings so that a leak would indicate, by oil or coolant, which ring was leaking.
Gardners used small O rings to seal coolant passages between block and head.
Other manufacturers , as a result of development, went into production with gaskets that were locally thickened with integral rings, or silicone rubber rings, to seal oil or coolant passages.
Quite a few engines, particularly air cooled ones, used a Wills ring to seal between barrel and head.
Push rod, or camshaft drive, bores/tubes, being in a low pressure region, were often sealed with O rings.
Depending upon which stage, prototype engines could differ quite markedly from what eventually went into production, as problems arose and were, hopefully, solved.
(Compression ratios, gaskets, gears, drive arrangements, fuel feeds, could all change between prototype and production engines.)
So a prototype could be quite different from what eventually went into volume production.
Howard