In a clock the wheel drives the pinion. In a reduction gear like yours the pinion drives the wheel so friction should be less of a problem. However the anti-backlash mechanism, if I remember rightly, consists of two wheels with a spring so that the teeth are staggered under spring pressure. This will create rubbing where a normal gear only has rolling motion between the teeth.
When I worked in a radio shop in around 1960 we used to put a thin smear of light grease on the teeth to reduce the friction created by the anti-backlash gear. Yes dust is a problem and needs to be kept out. I remember one set that a customer who had recently returned from Kenya brought in – it was full of giant spiders, beetles, and other nasties which had produced an HT short!
Russell.