An odd one, that behaviour. It should not happen on any steam-engine.
My first thought was simply from not using the bypass-valve on the feed-pump (if one is fitted); but that would obviously leave the boiler full when stopped. That’s not the case, you say.
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The water rises and falls in the glass – so the fitting’s passages are clear. So too, we hope, the firebox water walls, which can eventually choke if the feed water is hard. An unchanging water level is an ominous sign!
The change you see does not seem the surging we might expect in a locomotive under way, as the water sloshes back and forth, especially in a long boiler. The level will wobble up and down a bit but should be fairly well settled at steady speed on a good track, and of course stationary.
However, you describe the glass responding to the regulator being opened and closed.
That makes me ask:
1) Is this only when accelerating or decelerating (though it would have be rather brisk to make much difference) or when the locomotive is pulling at a steady speed as well?
2) Especially if the latter case, where are the steam take-off point and the gauge-glass top fitting, with respect to each other?
I don’t know the Modelworks ‘Britannia’ design and I am not sure of the full-size layout, but the main steam outlet should be high in the dome, whether the regulator is also there or in the smoke-box. Whereas the connections to the gauge-glass are normally on the backhead, and the other fittings take steam from a manifold above that. So all too far apart, in proportion to the boiler’s overall size, to affect each other.
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(You might ask, why not also put the gauge-glass top-fitting on the manifold. It’s not good practice because the flow of steam through valves to other fittings may, in theory at least, create sufficient depression in the gauge-glass connection to draw the water up in the glass, giving a false reading. How real the effect is, I would not care to say and it would be specific to the boiler and its operation anyway, but it is Something Not Done!)